Academic Background: PhD, University of Wisconsin, 1955; BS, Carleton College, 1951.
Research Interests: Protein folding, enzyme kinetics, biochemical mechanisms.
Appointments: Alumni Endowed Professor, Department of Biochemistry and Molecular Biophysics; Former Department Chair.
Awards: American Society for Biochemistry and Molecular Biology William C. Rose Award, 2000.
Publications: Over 200 peer-reviewed articles, including 'Protein Folding and Aggregation' (Wiley, 2010).